Research Projects

This page highlights a selection of the research, education, and outreach projects I have contributed to throughout my graduate career. Spanning fossil mammals, geochemistry, artificial intelligence, and STEM education, these projects reflect my interdisciplinary approach to understanding the past while building connections among science, technology, and society.

Fossil Horse Macroevolution

Investigating macroevolutionary transitions in Neogene Equidae using phenomics and machine learning

Fossil horses (Family Equidae) provide one of the most complete records of mammalian evolution in the fossil record, spanning over 55 million years in North America. My dissertation research focuses on macroevolutionary transitions in Neogene fossil horses, leveraging phenomics — the large-scale, quantitative measurement of phenotypic variation, alongside machine learning to detect patterns of evolutionary change across geographic and temporal scales. By integrating museum collections from across North America with computational approaches, I aim to reveal how populations of fossil horses changed morphologically across major climatic and environmental transitions.

Geological range and North American Land Mammal Ages of latest Hemphillian equid faunas
Figure 2. Geological ranges and North American Land Mammal Ages (NALMAs) of latest Hemphillian (Hh4) equid faunas in North America, including four widespread and two endemic species. (Killingsworth and MacFadden, 2023).

A central component of this work involves applying interpretable deep learning models to images of fossil horse teeth, which preserve detailed morphological information about diet, ecology, and phylogenetic relationships. This approach allows me to quantify subtle variation across large sample sizes that would be impractical to measure manually, opening new windows into paleopopulation structure and evolutionary dynamics. This research was selected for presentation in the Romer Prize Session at the 2026 Society of Vertebrate Paleontology annual meeting.

I also investigate species occurrence patterns among Late Miocene horses in Florida, examining whether gaps in the fossil record reflect genuine ecological absences, sampling biases, or both. This work addresses a fundamental challenge in interpreting evolutionary and biogeographic patterns from museum collections.

Publications

Killingsworth et al. ( )
MacFadden et al. ( , )

Acknowledgements

This research was supported by:

American Museum of Natural History, UC Museum of Paleontology, Smithsonian Institution, Florida Museum of Natural History